25.
Syngonium mauroanum Birdsey ex Bunting,
Baileya 14:18. 1966. TYPE: Panama (exact locality unknown, cultivated
at Amazon Gardens, Miami, Florida) Bunting 1441 (BH, holotype).
Juvenile plants with stems at first short, forming rosettes, becoming
elongate, green; petioles 6-13 cm long, sheathed to the middle or
somewhat above the middle, sharply 1-ribbed from the sheath to the
blade; blades sagittate, the primary lateral veins and many secondary
veins sunken above and beneath, the upper surface marked along the
midrib and sometimes along the larger lateral veins with a discolorous
band of gray or yellowish green, the anterior lobe ovate, acuminate,
5-16 cm long, the posterior lobes triangular to ovate, acute to
rounded at the apex, 2-10 cm long. Adult plants with stems not glaucous,
scandent, ap-pressed to trees; intemodes 1-12 cm long, 1-1.5 cm
wide; petioles sheathed 1/2-3/4 their length; blades thin, subtrisect
to usually trisect, rarely 5-segmented, medium green, matte above,
slightly paler and matte beneath; leaflets free or sometimes confluent;
median leaflet elliptic to ovate-elliptic, 12-21(-30) cm long, 3.5-10(-12)
cm wide, acuminate at the apex, obtuse to acute and attenuate at
the base, the margin minutely undulate and sometimes appearing toothed
on drying;
lateral leaflets inequilateral, sometimes auriculate, the auricle
usually conspicuously hastate, narrowly rounded at the apex; primary
lateral veins 6-10 pairs, distributed throughout the blade (but
thicker in the lower half of the blade), conspicuously sunken above,
raised beneath; principal, secondary, tertiary and some quaternary
veins sunken above, raised beneath, the reticulate veins clearly
visible beneath.
Inflorescences 1 or 2 per axil; peduncles 3.5-9 cm
long and erect at anthesis, 8-14 cm long and pendent in fruit; spathe
tube ellipsoid to narrowly ovoid, 3.5-5 cm long, 1.5-2.5 cm diam.,
green outside, dark violet purple inside;
spathe blade ovate-elliptic, 6-8 cm long, 4-5 cm wide, acuminate,
white; spadix 6.5-10 cm long, narrowly clavate; pistillate portion
of the spadix ca. 1 cm long on the back side, ca. 1.5 cm long on
the front side, the flowers irregularly rhombic at the apex, 2.3-3
mm diam., the stigma ca. 1.2 mm diam.; staminate portion of the
spadix white, the sterile part ca. 1.5 cm long, the fertile part
5-6 cm long, the synandrium irregularly rounded to rhombic at the
apex, the margin irregular, the apex not drying smooth, irregularly
undulate.
Infructescences 4.5-7.5 cm diam., red on the outside,
violet purple within; syncarp brown, 4.5-6 cm long, 2.5-4 cm diam.;
mesocarp white; seeds ovoid, 5-6 mm long, 4-5 mm diam., dark brown.
Figs. 52, 57, 58.
DISTRIBUTION: Syngonium mauroanum is known from Costa Rica
and Panama in seasonally dry parts of tropical moist forest and
especially from premon-tane moist forest life zones. It is an interesting
testimony to the apparent accuracy of the Holdridge Life Zone Maps
that the only collection from Costa Rica was also found in a narrow
band of premontane moist forest in Guanacaste and that the species
has been collected in the small isolated patch of premontane moist
forest around Santa Fe in Veraguas. The species ranges from sea
level to about 450 m elevation.
The species can be recognized by its usually hastate-lobed blades
with distinctive prominent venation. It is not easily confused with
any other species because of its relative isolation and narrow ecological
range, but the species is closest to S. standleyanum, a species
from premontane wet forest in Guanacaste Province, Costa Rica. That
species often shows no sign of auricles on the lateral leaflets
(a feature rarely exhibited by S. mauroanum) and has a densely
papillate stem. Syngonium standleyanum
may ultimately prove to be merely an ecological variant of this
species.
Flowers are known from April to June and immature fruits in November
and January.
COSTA RICA: PUNTARENAS: N Puntarenas, Janzen 10731
(MO); Ujarras de Buenos Aires, Pittier 11121 (US).
PANAMA: CHIRIQUI: Near Remedies, Woodson et al. 1189 (F, GH, MO).
HERRERA: S of Ocii, Lewis et al. 1637 (GH, US). LOS SANTOS: Vicinity
of Tonosi, Croat 9763 (MO), Lewis et al. 1561 (GH, K, US), Lewis
et al. 2193 (MO), Tyson et al. 2954 (MO, SCZ). PANAMA: Vicinity
of Panama City, Bartlett & Lasser 16708 (DUKE, MICH, MO). VERAGUAS:
Vicinity Santa Fe, Croat 27345, 34249 (MO); E edge of Escuela Agricola
Alto Piedra, Croat & Folsom 33981 (MO); Vicinity of Santa Fe,
Nee 8038 (GH, MO), Luteyn 4048 (DUKE, MO); S of Santiago, Nee 8231
(MO), Tyson 6045 (SCZ).
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